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* Lakshmanan & Parkash (1980) studied the nature and function of the layer of muscle fiber in the prepuce. The authors discovered the whorled pattern that produces a sphincter. They commented on the manner in which the [[foreskin]] is held close to the [[glans penis]]. They were the first to describe the [[gliding action]] of the foreskin that facilitates intercourse.<ref name="lakshmanan1980">{{REFjournal
|last= Lakshmanan
|init=S
|pages=134-7
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}}</ref>
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