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|accessdate=2022-10-07
}}</ref> Moistness also may be maintained by [[transudation]].<ref name="cold-taylor1999" />
Lakshmanan & Parkash (1980) described the muscle fibers in the prepuce (known as the [[dartos]], which contract to keep the prepuce snugly close to the glans penis. The muscle fibers form a whorl at the tip to keep the tip of the foreskin and the preputial sac closed against the outside world and contamination.<ref>{{REFjournal
|last=Lakshmanan
|first=
|init=S
|author-link=
|last2=Parkash
|first2=
|init2=S
|author2-link=
|etal=no
|title=Human prepuce: some aspects of structure and function
|trans-title=
|language=
|journal=Indian J Surg
|location=
|date=1980
|volume=44
|issue=
|article=
|page=
|pages=134-7
|url=http://www.cirp.org/library/anatomy/lakshmanan/
|archived=
|quote=
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|DOI=
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}}</ref>