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→Immunological functions: Add HIV information
|DOI=
|accessdate=2019-10-13
}}</ref> Fleiss ''et al''. noted the presence of Langerhans cells in the prepuce.<ref name="fleiss-hodges-vanhowe1998"/> De Witte ''et al''. (2007) have discovered that Langerhans cells produce Langerin, which is a "natural barrier to HIV infection".<ref>{{REFjournal |last=de Witte |first=Lot |first2=Alexey |last2=Nabatov |first3=Marjorie |last3=Pion |first4=Donna |last4=Fluitsma |first5=Marein A.W.P. |last5=de Jong |first6=Tanja |last6=de Gruijl |first7=Vincent |last7=Piguet |first8=Yvette |last8=van Kooyk |first9=Teunis B.H. |last9=Geijtenbeek |title=Langerin is a natural barrier to HIV-1 transmission by Langerhans cells |journal=Nature Medicine |volume= |issue= |pages= |url=https://www.nature.com/articles/nm1541 |quote= |pubmedID=17334373 |pubmedCID= |DOI=10.1038/nm1541 |date=2007-03-04 |accessdate=2011-06-28}}</ref>
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