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{{FleissP HodgesF VanHoweRS 1998}}
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Fleiss et al. (1998) reported that the moisture found under the foreskin contains pheromones such as androsterone and others.<ref name="fleiss-hodges-vanhowe1998">{{REFjournal |last=Fleiss |init=PM |author-link=Paul M. Fleiss |last2=Hodges |init2=FM |author2-link=Frederick M. Hodges |last3=Van Howe |init3=RS |author3-link=Robert S. Van Howe |title=Immunological functions of the human prepuce |journal=Sex Trans Infect |date=FleissP HodgesF VanHoweRS 1998-10 |volume=74 |issue=5 |pages=364-67 |url=https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1758142/pdf/v074p00364.pdf |quote= |pubmedID=10195034 |pubmedCID= |DOI= |accessdate=2021-06-11}}</ref>
Androsterone is found in the human axilla and [[skin]] as well as in the [[urine]]. It may also be secreted by human sebaceous glands. It is described as having a musky odor similar to that of androstenol. Androsterone has been found to affect human behavior when smelled.<ref>{{REFbook