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|date=2012
|accessdate=2022-10-07
}}</ref> Moistness also may be maintained by [[transudation]].<ref name="cold-taylor1999" />The preputial sac may also receive [[pre-ejaculate]].
Lakshmanan & Parkash (1980) described the muscle fibers in the [[prepuce]] (known as the [[dartos]]), which contract to keep the prepuce snugly close to the [[glans penis]]. The muscle fibers form a whorl at the tip to keep the tip of the [[foreskin]] and keep the preputial sac closed against the outside world and contamination.<ref>{{REFjournal