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* Parkash et al (1973)studied the nature of the material which collects in the [[preputial sac]] of [[intact]] males. It was determined to be cast off "epithelial debris".<ref name="parkash1973">{{REFjournal |last=Parkash |first= |init=S |author-link= |last2=Jeyakumar |first2= |init2=S |author2-link= |last3=Subramanyan |first3= |init3=K |author3-link= |last4=Chaudhuri |first4= |init4=S |author4-link= |etal=no |title=Human subpreputial collection: its nature and formation |trans-title= |language= |journal=J Urol |location= |date=1973-08 |volume=110 |issue=2 |pages=211-2 |url=https://www.cirp.org/library/anatomy/parkash/ |archived= |quote=A detailed study of the subpreputial collections and histological study of 128 specimens led us to conclude that there appears to be no evidence of any glandular tissue in the subpreputial region of the penis. Smegma is an epithelial debris collecting in this space. |pubmedID=4722614 |pubmedCID= |DOI=10.1016/s0022-5347(17)60164-2 |accessdate=2024-07-05}}</ref>
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